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Biomedical Sciences

All courses, faculty listings, and curricular and degree requirements described herein are subject to change or deletion without notice.

Professors

Ruben Abagyan, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Cristian Achim, MD, PhD, Psychiatry and Neuropathology (Adjunct)

Susan Ackerman, PhD, Cellular and Molecular Medicine

Joseph Adams, PhD, Pharmacology

Rommie Amaro, PhD, Chemistry and Biochemistry

Corina Antal, PhD, Pharmacology

Radha Ayyagari, PhD, Radiology/Ophthalmology

Maksim Bazhenov, PhD, Medicine

Bogdan Bintu, PhD, Cellular and Molecular Medicine

Gerry R. Boss, MD, Medicine

Nunzio Bottini, MD, PhD, Medicine

Joan Heller Brown, PhD, Pharmacology (Chair)

Laurence L. Brunton, PhD, Pharmacology/Medicine (Emeritus)

Jack Bui, MD, PhD, Pathology

Michael Burkart, PhD, Professor, Chemistry and Biochemistry

Geoffrey Chang, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

John Chang, MD, Medicine

Fabian Rivera-Chávez, PhD, Pediatrics

Ju Chen, PhD, Medicine (In Residence)

David Cheresh, PhD, Pathology

Neil Chi, MD, PhD, Medicine

Jerold J. M. Chun, PhD, Pharmacology (Adjunct)

Don W. Cleveland, PhD, Cellular and Molecular Medicine (Chair)/Medicine

Nicole Coufal, MD, PhD, Pediatrics (Adjunct)

Leslie A. Crews, PhD, Medicine

Michael Croft, PhD, Medicine

Shane Crotty, PhD, Medicine (Adjunct)

Antonio De Maio, PhD, Surgery

Edward Dennis, PhD, Pharmacology/Chemistry and Biochemistry

Arshad Desai, PhD, Cellular and Molecular Medicine

Neal Devaraj, PhD, Chemistry and Biochemistry

Jack E. Dixon, PhD, Pharmacology/Cellular and Molecular Medicine/Chemistry and Biochemistry (Dean, Scientific Affairs)

Daniel Donoghue, PhD, Chemistry and Biochemistry

Pieter Dorrestein, PhD, Pharmacy/Chemistry and Biochemistry

Steven F. Dowdy, PhD, Cellular and Molecular Medicine

Tal Einav, PhD, La Jolla Institute of Immunology

Mark H. Ellisman, PhD, Neurosciences

Adam Engler, PhD, Bioengineering/UC San Diego Stem Cell Institute

Peter Ernst, DVM, PhD, Pathology

Jeffrey Esko, PhD, Cellular and Molecular Medicine

Ronald M. Evans, PhD, Biology (Adjunct)

Sylvia Evans, PhD, Pharmacy/Pharmacology

Xi Fang, PhD, Medicine

Gen-Sheng Feng, PhD, Pathology

Cole Ferguson, MD, PhD, Pathology

Fleur M. Ferguson, PhD, Chemistry and Biochemistry/Skaggs School of Pharmacy and Pharmaceutical Sciences

Susan Ferro-Novick, PhD, Cellular and Molecular Medicine

Kelly Frazer, PhD, Pediatrics

Theodore Friedmann, MD, Pediatrics

Frank Funari, PhD, Pathology

Pascal Gagneux, PhD, Cellular and Molecular Medicine (Adjunct)

Richard Gallo, MD, PhD, Medicine

William Gerwick, PhD, Oceanography/Skaggs School of Pharmacy and Pharmaceutical Sciences

Partho Ghosh, PhD, Chemistry and Biochemistry

Pradipta Ghosh, MD, Medicine

Mark Ginsberg, MD, Medicine

Christopher K. Glass, MD, PhD, Cellular and Molecular Medicine/Medicine

Joseph G. Gleeson, MD, Neurosciences

Ananda Goldrath, PhD, Molecular Biology

Steven Gonias, MD, PhD, Pathology (Chair)

Philip Gordts, PhD, Department of Medicine

John C. Guatelli, PhD, Medicine

Shiri Gur-Cohen, PhD, Medicine

Asa Gustafsson, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Silvio Gutkind, PhD, Pharmacology

Gabriel Haddad, MD, Pediatrics (Chair)/Neurosciences

Bruce Hamilton, PhD, Cellular and Molecular Medicine/Medicine

Tracy Handel, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Stephen Hedrick, PhD, Cellular and Molecular Medicine

Sven Heinz, PhD, Medicine

Robert Hevner, MD, PhD, Pathology

Michael C. Hogan, PhD, Medicine (Adjunct)

Vivian Hook, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Anthony Hunter, PhD, Molecular Biology

Trey Ideker, PhD, Medicine

Catriona Jamieson, MD, PhD, Medicine

Yishi Jin, PhD, Cellular and Molecular Medicine/Biological Sciences

Michael Karin, PhD, Pharmacology

Alexander Kauffman, PhD, Reproductive Medicine

Dan Kaufman, MD, PhD, Medicine

John Kelsoe, MD, Psychiatry

David Kleinfeld, PhD, Physics

Rob Knight, PhD, Pediatrics

Richard Kolodner, PhD, Cellular and Molecular Medicine

Elizabeth A. Komives, PhD, Chemistry and Biochemistry

Mitchell Kronenberg, PhD, Molecular Biology

Mark Lawson, PhD, Reproductive Medicine

Hyam L. Leffert, MD, Pharmacology

Andres Leschziner, PhD, Cellular and Molecular Medicine

Willis Li, PhD, Medicine

George Liu, MD, PhD, Pediatrics

James McKerrow, MD, PhD, Dean, Skaggs School of Pharmacy and Pharmaceutical Sciences

Hiruy Meharena, PhD, Neurobiology

Jill Mesirov, PhD, Medicine

Yury Miller, MD, PhD, Medicine

Bradley Moore, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Alysson Muotri, PhD, Pediatrics/Cellular and Molecular Medicine

Cornelis Murre, PhD, Molecular Biology

Samuel Myers, PhD, La Jolla Institute for Immunology

Alexandra Newton, PhD, Pharmacology

Victor Nizet, MD, Pediatrics and Skaggs School of Pharmacy and Pharmaceutical Sciences

Peter Novick, PhD, Cellular and Molecular Medicine

Karen Oegema, PhD, Cellular and Molecular Medicine

Abraham Palmer, PhD, Psychiatry

Mana Parast, MD, PhD, Pathology

Hemal Patel, PhD, Anesthesiology

Renate B. Pilz, MD, Medicine (In Residence)

Frank L. Powell Jr., PhD, Medicine

Oswald Quehenberger, PhD, Medicine

Manuela Raffatellu, MD, Pediatrics

Tariq Rana, PhD, Pediatrics (Adjunct)

Anjana Rao, PhD, Pharmacology (Adjunct)

Samara L. Reck-Peterson, PhD, Cellular and Molecular Medicine

Bing Ren, PhD, Cellular and Molecular Medicine

Michael G. Rosenfeld, MD, Medicine

Robert Ross, MD, Medicine (In Residence)

Subhojit Roy, MD, PhD, Pathology

Alan Saltiel, PhD, Medicine

Guy Salvesen, PhD, Pathology (Adjunct)

Maike Sander, MD, Pediatrics/Cellular and Molecular Medicine

Richard Scheuermann, PhD, Pathology

David Schlaepfer, PhD, Reproductive Medicine

Geert Schmid-Schoenbein, PhD, Bioengineering

Stephen Schoenberger, PhD, Medicine

Johannes Schoeneberg, PhD, Pharmacology

Nicholas Schork, PhD, Psychiatry and Biostatistics

Jonathan Sebat, PhD, Psychiatry/Cellular and Molecular Medicine

Jianhua Shao, MD, PhD, Pediatrics

Sanford Shattil, MD, Medicine

Reuben Shaw, PhD, Molecular Biology

Aleem Siddiqui, PhD, Medicine

Dionicio Siegel, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Christina Sigurdson, DVM, PhD, Pathology

Monique Smith, PhD, Neurobiology

Evan Snyder, MD, PhD, Pediatrics (Associate Physician)

Francesca Soncin, PhD, Pathology

Deborah Spector, PhD, Cellular and Molecular Medicine

Nicole Steinmetz, PhD, NanoEngineering

Ina Stelzer, PhD, Pathology

Xin Sun, PhD, Pediatrics

Roger Sunahara, PhD, Pharmacology

Palmer W. Taylor, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Susan S. Taylor, PhD, Chemistry and Biochemistry, Pharmacology

David Traver, PhD, Cellular and Molecular Medicine

JoAnn Trejo, PhD, Pharmacology

Emily Troemel, PhD, Cell and Developmental Biology

Robert H. Tukey, PhD, Pharmacology/Chemistry and Biochemistry

Ajit P. Varki, MD, Medicine

Judith A. Varner, PhD, Pathology

Francisco Villarreal, MD, PhD, Medicine (Adjunct)

Dong Wang, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Emily Wang, PhD, Pathology

Jean Wang, PhD, Medicine

Nicholas J. G. Webster, PhD, Medicine (In Residence)

Miles Wilkinson, PhD, Reproductive Medicine

Karl Willert, PhD, Cellular and Molecular Medicine

Elizabeth Winzeler, PhD, Pediatrics

Joseph L. Witzum, MD, Medicine

Tony L. Yaksh, PhD, Anesthesiology/Pharmacology

Jing Yang, PhD, Pharmacology/Pediatrics

Gene Yeo, PhD, Cellular and Molecular Medicine

Wei Ying, PhD, Medicine

Maurizio Zanetti, MD, Medicine (Emeritus)

Karsten Zengler, PhD, Pediatrics

Dong-Er Zhang, PhD, Pathology/Biological Sciences

Jin Zhang, PhD, Pharmacology

Kun Zhang, PhD, Bioengineering

Binhai Zheng, PhD, Neurosciences

Huilin Zhou, PhD, Cellular and Molecular Medicine

Yimin Zou, PhD, Neurobiology

Elina Zuniga, PhD, Molecular Biology

Associate Professors

Fotis Asimakopoulos, MD, PhD, Medicine

Ferhat Ay, PhD, Pediatrics (Adjunct)

Carlo Ballatore, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Rafael Bejar, MD, PhD, Internal Medicine

Alejandro (Alex) Chavez, PhD, Pediatrics

Stephanie Cherqui, PhD, Pediatrics (In Residence)

Cosimo Commisso, PhD, Surgery

Kevin Corbett, PhD, Cellular and Molecular Medicine

Ben Croker, PhD, Pediatrics

Kamil Godula, PhD, Chemistry and Biochemistry

Thomas Hnasko, PhD, Neurosciences

Lilia Iakoucheva, PhD, Psychiatry

Juliana Idoyaga, PhD, Pharmacology and Molecular Biology

Mohit Jain, MD, PhD, Medicine, Pharmacology 

William Joiner, PhD, Pharmacology

Irina Kufareva, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Nathan Lewis, PhD, Pediatrics

Li-Fan Lu, PhD, Molecular Biology

Ayako Makino, PhD, Medicine

David McCulley, MD, Pediatrics

Jerome Mertens, PhD, Neurosciences

Anthony Molina, PhD, Medicine

Gerald P. Morris, MD, PhD, Pathology

Larissa Podust, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Christopher Putnam, PhD, Medicine

Zoran Radic, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Robert Rissman, PhD, Neuroscience

Simon Schenk, PhD, Orthopaedic Surgery

George Sen, PhD, Medicine/Cellular and Molecular Medicine

Farah Sheikh, PhD, Medicine (Adjunct)

Dwayne Stupack, PhD, Reproductive Medicine

Pandurangan Vijayanand, MD, PhD, Medicine

Jill Wildonger, PhD, Pediatrics

Peter Zage, MD, PhD, Pediatrics

Sheng Zhong, PhD, Bioengineering

Assistant Professors

Ludmil Alexandrov, PhD, Cellular and Molecular Medicine

Christopher Benner, PhD, Medicine

Zea Borok, PhD, Medicine

Aaron Carlin, MD, PhD, Medicine

Yuan Chen, PhD, Surgery

Hiuting Chu, PhD, Pathology

Heidi Cook-Andersen, MD, PhD, Reproductive Medicine and Biological Sciences

Richard Daneman, PhD, Pharmacology and Neuroscience

Matthew Daugherty, PhD, Molecular Biology

Emma Farley, PhD, Medicine, Molecular Biology

Stephanie Fraley, PhD, Bioengineering

Kyle Gaulton, PhD, Pediatrics

Jack A. Gilbert, PhD, Pediatrics

David Gonzalez, PhD, Pharmacology

Alon Goren, PhD, Medicine

Melissa Gymrek, PhD, Medicine

Matt Hangauer, PhD, Dermatology

Ryan Hibbs, PhD, Neurobiology

Wendy Huang, PhD, Cellular and Molecular Medicine

Enfu Hui, PhD, Cell and Developmental Biology

Louise Laurent, MD, PhD, Obstetrics, Gynecology, and Reproductive Sciences

Amanda L. Lewis, PhD, Obstetrics, Gynecology, and Reproductive Sciences

Byungkook Lim, PhD, Neurobiology

Amit Majithia, MD, Medicine and Pediatrics

Prashant Mali, PhD, Bioengineering

Karen Mestan, MD, Pediatrics

Eran Mukamel, PhD, Cognitive Science

Anthony O’Donoghue, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences

Erica Ollmann Saphire, PhD, Medicine (Adjunct)

Andrew Sharabi, MD, PhD, Clinical Radiation Medicine and Applied Science

Robert Signer, PhD, Medicine

Tatum Simonson, PhD, Medicine

Scott Sternson, PhD, Neurosciences

Bryan Sun, MD, PhD, Dermatology

Francesca Telese, PhD, Medicine

Elizabeth Villa, PhD, Chemistry and Biochemistry

Karl Wahlin, PhD, Opthalmology

Samuel R. Ward, PhD, Orthopaedic Surgery and Radiology

Daniela Weiskopf, PhD, Medicine (Adjunct)

Amir Zarrinpar, MD, PhD, Medicine