Biomedical Sciences
All courses, faculty listings, and curricular and degree requirements described herein are subject to change or deletion without notice.
Professors
Ruben Abagyan, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Cristian Achim, MD, PhD, Psychiatry and Neuropathology (Adjunct)
Susan Ackerman, PhD, Cellular and Molecular Medicine
Joseph Adams, PhD, Pharmacology
Rommie Amaro, PhD, Chemistry and Biochemistry
Corina Antal, PhD, Pharmacology
Radha Ayyagari, PhD, Radiology/Ophthalmology
Maksim Bazhenov, PhD, Medicine
Bogdan Bintu, PhD, Cellular and Molecular Medicine
Gerry R. Boss, MD, Medicine
Nunzio Bottini, MD, PhD, Medicine
Joan Heller Brown, PhD, Pharmacology (Chair)
Laurence L. Brunton, PhD, Pharmacology/Medicine (Emeritus)
Jack Bui, MD, PhD, Pathology
Michael Burkart, PhD, Professor, Chemistry and Biochemistry
Geoffrey Chang, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
John Chang, MD, Medicine
Fabian Rivera-Chávez, PhD, Pediatrics
Ju Chen, PhD, Medicine (In Residence)
David Cheresh, PhD, Pathology
Neil Chi, MD, PhD, Medicine
Jerold J. M. Chun, PhD, Pharmacology (Adjunct)
Don W. Cleveland, PhD, Cellular and Molecular Medicine (Chair)/Medicine
Nicole Coufal, MD, PhD, Pediatrics (Adjunct)
Leslie A. Crews, PhD, Medicine
Michael Croft, PhD, Medicine
Shane Crotty, PhD, Medicine (Adjunct)
Antonio De Maio, PhD, Surgery
Edward Dennis, PhD, Pharmacology/Chemistry and Biochemistry
Arshad Desai, PhD, Cellular and Molecular Medicine
Neal Devaraj, PhD, Chemistry and Biochemistry
Jack E. Dixon, PhD, Pharmacology/Cellular and Molecular Medicine/Chemistry and Biochemistry (Dean, Scientific Affairs)
Daniel Donoghue, PhD, Chemistry and Biochemistry
Pieter Dorrestein, PhD, Pharmacy/Chemistry and Biochemistry
Steven F. Dowdy, PhD, Cellular and Molecular Medicine
Tal Einav, PhD, La Jolla Institute of Immunology
Mark H. Ellisman, PhD, Neurosciences
Adam Engler, PhD, Bioengineering/UC San Diego Stem Cell Institute
Peter Ernst, DVM, PhD, Pathology
Jeffrey Esko, PhD, Cellular and Molecular Medicine
Ronald M. Evans, PhD, Biology (Adjunct)
Sylvia Evans, PhD, Pharmacy/Pharmacology
Xi Fang, PhD, Medicine
Gen-Sheng Feng, PhD, Pathology
Cole Ferguson, MD, PhD, Pathology
Fleur M. Ferguson, PhD, Chemistry and Biochemistry/Skaggs School of Pharmacy and Pharmaceutical Sciences
Susan Ferro-Novick, PhD, Cellular and Molecular Medicine
Kelly Frazer, PhD, Pediatrics
Theodore Friedmann, MD, Pediatrics
Frank Funari, PhD, Pathology
Pascal Gagneux, PhD, Cellular and Molecular Medicine (Adjunct)
Richard Gallo, MD, PhD, Medicine
William Gerwick, PhD, Oceanography/Skaggs School of Pharmacy and Pharmaceutical Sciences
Partho Ghosh, PhD, Chemistry and Biochemistry
Pradipta Ghosh, MD, Medicine
Mark Ginsberg, MD, Medicine
Christopher K. Glass, MD, PhD, Cellular and Molecular Medicine/Medicine
Joseph G. Gleeson, MD, Neurosciences
Ananda Goldrath, PhD, Molecular Biology
Steven Gonias, MD, PhD, Pathology (Chair)
Philip Gordts, PhD, Department of Medicine
John C. Guatelli, PhD, Medicine
Shiri Gur-Cohen, PhD, Medicine
Asa Gustafsson, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Silvio Gutkind, PhD, Pharmacology
Gabriel Haddad, MD, Pediatrics (Chair)/Neurosciences
Bruce Hamilton, PhD, Cellular and Molecular Medicine/Medicine
Tracy Handel, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Stephen Hedrick, PhD, Cellular and Molecular Medicine
Sven Heinz, PhD, Medicine
Robert Hevner, MD, PhD, Pathology
Michael C. Hogan, PhD, Medicine (Adjunct)
Vivian Hook, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Anthony Hunter, PhD, Molecular Biology
Trey Ideker, PhD, Medicine
Catriona Jamieson, MD, PhD, Medicine
Yishi Jin, PhD, Cellular and Molecular Medicine/Biological Sciences
Michael Karin, PhD, Pharmacology
Alexander Kauffman, PhD, Reproductive Medicine
Dan Kaufman, MD, PhD, Medicine
John Kelsoe, MD, Psychiatry
David Kleinfeld, PhD, Physics
Rob Knight, PhD, Pediatrics
Richard Kolodner, PhD, Cellular and Molecular Medicine
Elizabeth A. Komives, PhD, Chemistry and Biochemistry
Mitchell Kronenberg, PhD, Molecular Biology
Mark Lawson, PhD, Reproductive Medicine
Hyam L. Leffert, MD, Pharmacology
Andres Leschziner, PhD, Cellular and Molecular Medicine
Willis Li, PhD, Medicine
George Liu, MD, PhD, Pediatrics
James McKerrow, MD, PhD, Dean, Skaggs School of Pharmacy and Pharmaceutical Sciences
Hiruy Meharena, PhD, Neurobiology
Jill Mesirov, PhD, Medicine
Yury Miller, MD, PhD, Medicine
Bradley Moore, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Alysson Muotri, PhD, Pediatrics/Cellular and Molecular Medicine
Cornelis Murre, PhD, Molecular Biology
Samuel Myers, PhD, La Jolla Institute for Immunology
Alexandra Newton, PhD, Pharmacology
Victor Nizet, MD, Pediatrics and Skaggs School of Pharmacy and Pharmaceutical Sciences
Peter Novick, PhD, Cellular and Molecular Medicine
Karen Oegema, PhD, Cellular and Molecular Medicine
Abraham Palmer, PhD, Psychiatry
Mana Parast, MD, PhD, Pathology
Hemal Patel, PhD, Anesthesiology
Renate B. Pilz, MD, Medicine (In Residence)
Frank L. Powell Jr., PhD, Medicine
Oswald Quehenberger, PhD, Medicine
Manuela Raffatellu, MD, Pediatrics
Tariq Rana, PhD, Pediatrics (Adjunct)
Anjana Rao, PhD, Pharmacology (Adjunct)
Samara L. Reck-Peterson, PhD, Cellular and Molecular Medicine
Bing Ren, PhD, Cellular and Molecular Medicine
Michael G. Rosenfeld, MD, Medicine
Robert Ross, MD, Medicine (In Residence)
Subhojit Roy, MD, PhD, Pathology
Alan Saltiel, PhD, Medicine
Guy Salvesen, PhD, Pathology (Adjunct)
Maike Sander, MD, Pediatrics/Cellular and Molecular Medicine
Richard Scheuermann, PhD, Pathology
David Schlaepfer, PhD, Reproductive Medicine
Geert Schmid-Schoenbein, PhD, Bioengineering
Stephen Schoenberger, PhD, Medicine
Johannes Schoeneberg, PhD, Pharmacology
Nicholas Schork, PhD, Psychiatry and Biostatistics
Jonathan Sebat, PhD, Psychiatry/Cellular and Molecular Medicine
Jianhua Shao, MD, PhD, Pediatrics
Sanford Shattil, MD, Medicine
Reuben Shaw, PhD, Molecular Biology
Aleem Siddiqui, PhD, Medicine
Dionicio Siegel, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Christina Sigurdson, DVM, PhD, Pathology
Monique Smith, PhD, Neurobiology
Evan Snyder, MD, PhD, Pediatrics (Associate Physician)
Francesca Soncin, PhD, Pathology
Deborah Spector, PhD, Cellular and Molecular Medicine
Nicole Steinmetz, PhD, NanoEngineering
Ina Stelzer, PhD, Pathology
Xin Sun, PhD, Pediatrics
Roger Sunahara, PhD, Pharmacology
Palmer W. Taylor, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Susan S. Taylor, PhD, Chemistry and Biochemistry, Pharmacology
David Traver, PhD, Cellular and Molecular Medicine
JoAnn Trejo, PhD, Pharmacology
Emily Troemel, PhD, Cell and Developmental Biology
Robert H. Tukey, PhD, Pharmacology/Chemistry and Biochemistry
Ajit P. Varki, MD, Medicine
Judith A. Varner, PhD, Pathology
Francisco Villarreal, MD, PhD, Medicine (Adjunct)
Dong Wang, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Emily Wang, PhD, Pathology
Jean Wang, PhD, Medicine
Nicholas J. G. Webster, PhD, Medicine (In Residence)
Miles Wilkinson, PhD, Reproductive Medicine
Karl Willert, PhD, Cellular and Molecular Medicine
Elizabeth Winzeler, PhD, Pediatrics
Joseph L. Witzum, MD, Medicine
Tony L. Yaksh, PhD, Anesthesiology/Pharmacology
Jing Yang, PhD, Pharmacology/Pediatrics
Gene Yeo, PhD, Cellular and Molecular Medicine
Wei Ying, PhD, Medicine
Maurizio Zanetti, MD, Medicine (Emeritus)
Karsten Zengler, PhD, Pediatrics
Dong-Er Zhang, PhD, Pathology/Biological Sciences
Jin Zhang, PhD, Pharmacology
Kun Zhang, PhD, Bioengineering
Binhai Zheng, PhD, Neurosciences
Huilin Zhou, PhD, Cellular and Molecular Medicine
Yimin Zou, PhD, Neurobiology
Elina Zuniga, PhD, Molecular Biology
Associate Professors
Fotis Asimakopoulos, MD, PhD, Medicine
Ferhat Ay, PhD, Pediatrics (Adjunct)
Carlo Ballatore, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Rafael Bejar, MD, PhD, Internal Medicine
Alejandro (Alex) Chavez, PhD, Pediatrics
Stephanie Cherqui, PhD, Pediatrics (In Residence)
Cosimo Commisso, PhD, Surgery
Kevin Corbett, PhD, Cellular and Molecular Medicine
Ben Croker, PhD, Pediatrics
Kamil Godula, PhD, Chemistry and Biochemistry
Thomas Hnasko, PhD, Neurosciences
Lilia Iakoucheva, PhD, Psychiatry
Juliana Idoyaga, PhD, Pharmacology and Molecular Biology
Mohit Jain, MD, PhD, Medicine, Pharmacology
William Joiner, PhD, Pharmacology
Irina Kufareva, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Nathan Lewis, PhD, Pediatrics
Li-Fan Lu, PhD, Molecular Biology
Ayako Makino, PhD, Medicine
David McCulley, MD, Pediatrics
Jerome Mertens, PhD, Neurosciences
Anthony Molina, PhD, Medicine
Gerald P. Morris, MD, PhD, Pathology
Larissa Podust, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Christopher Putnam, PhD, Medicine
Zoran Radic, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Robert Rissman, PhD, Neuroscience
Simon Schenk, PhD, Orthopaedic Surgery
George Sen, PhD, Medicine/Cellular and Molecular Medicine
Farah Sheikh, PhD, Medicine (Adjunct)
Dwayne Stupack, PhD, Reproductive Medicine
Pandurangan Vijayanand, MD, PhD, Medicine
Jill Wildonger, PhD, Pediatrics
Peter Zage, MD, PhD, Pediatrics
Sheng Zhong, PhD, Bioengineering
Assistant Professors
Ludmil Alexandrov, PhD, Cellular and Molecular Medicine
Christopher Benner, PhD, Medicine
Zea Borok, PhD, Medicine
Aaron Carlin, MD, PhD, Medicine
Yuan Chen, PhD, Surgery
Hiuting Chu, PhD, Pathology
Heidi Cook-Andersen, MD, PhD, Reproductive Medicine and Biological Sciences
Richard Daneman, PhD, Pharmacology and Neuroscience
Matthew Daugherty, PhD, Molecular Biology
Emma Farley, PhD, Medicine, Molecular Biology
Stephanie Fraley, PhD, Bioengineering
Kyle Gaulton, PhD, Pediatrics
Jack A. Gilbert, PhD, Pediatrics
David Gonzalez, PhD, Pharmacology
Alon Goren, PhD, Medicine
Melissa Gymrek, PhD, Medicine
Matt Hangauer, PhD, Dermatology
Ryan Hibbs, PhD, Neurobiology
Wendy Huang, PhD, Cellular and Molecular Medicine
Enfu Hui, PhD, Cell and Developmental Biology
Louise Laurent, MD, PhD, Obstetrics, Gynecology, and Reproductive Sciences
Amanda L. Lewis, PhD, Obstetrics, Gynecology, and Reproductive Sciences
Byungkook Lim, PhD, Neurobiology
Amit Majithia, MD, Medicine and Pediatrics
Prashant Mali, PhD, Bioengineering
Karen Mestan, MD, Pediatrics
Eran Mukamel, PhD, Cognitive Science
Anthony O’Donoghue, PhD, Skaggs School of Pharmacy and Pharmaceutical Sciences
Erica Ollmann Saphire, PhD, Medicine (Adjunct)
Andrew Sharabi, MD, PhD, Clinical Radiation Medicine and Applied Science
Robert Signer, PhD, Medicine
Tatum Simonson, PhD, Medicine
Scott Sternson, PhD, Neurosciences
Bryan Sun, MD, PhD, Dermatology
Francesca Telese, PhD, Medicine
Elizabeth Villa, PhD, Chemistry and Biochemistry
Karl Wahlin, PhD, Opthalmology
Samuel R. Ward, PhD, Orthopaedic Surgery and Radiology
Daniela Weiskopf, PhD, Medicine (Adjunct)
Amir Zarrinpar, MD, PhD, Medicine